Book Review: Developmental and Structural

Microscopic Venation Patterns of Leaves and their Importance in the Distinction of (Tropical) Species. Handbuch der Pflanzenanatomie XIV, part 4 Roth, Ingrid, 1996. ISBN 3-443-14023-8 (cloth US$98) 196 pp. Gebruder Borntraeger, Johannesstr. 3 A, D-70176 Stuttgart, Germany
- This book is so narrowly focused that it is hard to imagine the audience for it. It contains photographs and brief (usually) qualitative descriptions of the fine venation (without any, or hardly any, reference to the major venation or other leaf architectural features) of cleared and stained quarter centimeter squares cut from leaves of 170 species of dicot trees that happened to be growing in a single Venezuelan forest when it was harvested for an inventory 30 years ago.

There is little or no sense of intraspecific variability and extremely little synthesis. Keys are provided for the up to a dozen species in the larger families, but these wouldn't be particularly useful for identification, even elsewhere in Venezuela. Each of these families has many more species in northeastern South America and 1000-10,000 species worldwide. For the physiological ecologist, we have the already established observation, without quantitative testing, that venation gets denser and more compact as leaves get smaller with increasing stature in the forest. This applies, as well, to juveniles of canopy species, which also have less organized venation than adults of the same species.

Except for some casual citations in the introductory parts, there is virtually no acknowledgment of the extensive literature on leaf venation, particularly the rich paleobotanical and paleobotanically inspired literature. There isn't even any meaningful cross-referencing of the author's other books on the same trees. All told, then, this in not Prof. Roth's most interesting book.

It is easy to think of different approaches to the material that would have been far more useful and had a bigger impact than the approach taken here. Since more than 65,000 trees were felled for the original sample, there was a real opportunity to assess patterns of intraspecific variability, a potential problem that has never been dealt with in depth. Alternatively, Roth might have taken a more ecological tack and covered the broad range of environments in Venezuela that she has investigated over the years, with a serious attempt to test for environmental correlates of fine venation patterns. Finally, selecting a single family, or a few families, for more detailed comparative study would also have more clearly matched the title of this book than its actual contents. - James E. Eckenwalder, Department of Botany, University of Toronto.

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